Muslim brothers carried out two of the hotel suicide blasts in Sri Lanka on Easter Sunday, part of a wave of bombings that killed more than 320 people, police sources told AFP Tuesday.

The sons of a wealthy Colombo spice trader were among suicide bombers who hit three churches and three luxury hotels, investigators said.

An attack on a fourth hotel failed and helped lead police to the Islamist group now blamed for the assault, they added.

The brothers, whose names have not been revealed, were in their late twenties and operated their own "family cell", an investigation officer said.

The pair were key members of the Islamist National Thowheeth Jama'ath (NTJ) group which the government has blamed for the attacks, the official added.

The Islamic State group has claimed responsibility for the attacks, but the government has blamed the NTJ. A minister said Tuesday the bombers may have struck in revenge for attacks on two New Zealand mosques last month which left 50 dead.

One brother checked into the Cinnamon Grand hotel and the other the Shangri-La on Saturday.

The next morning, at virtually the same time, they went to the hotels' Easter Sunday breakfast buffets and blew up explosives-laden backpacks, the officer said.

Another bomb tore through a restaurant at the nearby Kingsbury hotel. Minutes before, similar explosions devastated three churches. Investigators said it was not known whether the brothers were in contact with the other bombers.

Another would-be suicide bomber walked into a fourth luxury hotel in Colombo on Sunday, official sources told AFP.

"This man had also checked into the hotel the previous day," the source said.

Fourth attacker

It was not known if his explosives failed or he had a change of heart.

But after the Shangri-la blast, staff at the unnamed hotel became suspicious and the man was tracked to a lodging near the capital. He blew himself up there when confronted by police, the source said. Two bystanders were also killed.

"What we have seen from the CCTV footage is that all the suicide bombers were carrying very heavy backpacks. These appear to be crude devices made locally," the source said.

With 321 people confirmed dead, including at least 39 foreign nationals, and over 500 wounded, Sri Lanka has declared a state of emergency and launched a desperate hunt to head off more attacks.

At least 40 people were under arrest Tuesday over one of the worst militant strikes ever staged in Asia.

The whereabouts of the brothers' parents was unknown. But the blasts had a further impact on the family.

One brother gave false identity details when he checked into the hotel, the investigator said. The other gave a real address which led police commandos to their family home in a commercial area of Colombo.

Children killed

"When the Special Task Force went there to investigate, one brother's wife set off explosives killing herself and her two children," the officer said.

"It was a single terror cell operated by one family," the investigator said.

"They had the cash and the motivation. They operated the cell and it is believed they influenced their extended family."

Three police commandos were killed in the blast, and several extended family members are among those in detention.

The brothers had been involved in their father's lucrative spice export business, investigators said.

A focus of the inquiry will be to find out whether there was a foreign influence in their radicalisation and how the children of such a wealthy family had become involved, an official source said.

"What we have gathered so far is that they had indicated to their close family what they were going to do," another senior police officer said.

"It looks like they were inspired by foreign terrorist groups, but to what extent they had direct links is still unclear."

Police alert

Sri Lanka police chief Pujith Jayasundara issued an alert on April 11 that the NTJ could launch suicide attacks against churches and other targets.

The whereabouts of the NTJ leader, Zahran Hashmi, is also unknown.

He was linked to the vandalising of Buddha statues on December 26 at the central town of Mawanella.

The local Muslim community had been complaining to authorities about Hashmi since 2017.

Residents of the village in the east of the country where he lived had demanded police action against him over his radical comments and acts, community leaders told AFP.

"He was a threat to moderate Muslims in the east and we had made several complaints," one Muslim leader told AFP.

The police chief's warnings about the NTJ were not passed to top ministers. A separate investigation is underway into why more was not done to stop the brothers and the other attackers.